Through the analysis of starch, phytoliths, and fungal remains in the pottery residues, we can now discuss several issues, including the function of pottery, the utilization of plant-based food resources, agricultural practices, ritual behaviors, social stratification, and regional interactions.
4.1. Evidence of fermentation method
Considering the presence of starch grains, phytoliths, and fungal remains in the pottery, it can be inferred that some vessels were used for brewing or drinking fermented beverages, while others served as cooking utensils or storage containers. Evidence of the pottery's contact with fermented food primarily comes from molds, yeast cells, and starch granules exhibiting fermentation characteristics. Among the 12 vessels examined, except for two (the ding caldron and pen basin), ten displayed a combination of cereals associated with yeast cells and/or molds. These observations suggest the use of the qu method for producing fermented beverages, with the primary fermentation agent consisting predominantly of Monascus, along with smaller amounts of Aspergillus and Rhizopus/Mucor. The raw brewing materials likely included rice, millets, Job’s tears, wheat, snake gourd root, lily, and water chestnut (Fig. 6; Figure S2).
The use of Monascus mold and rice to create the qu starter compound for making red rice beer represents the oldest brewing technique in China, dating back to the Shangshan culture in the lower Yangtze River region around 9000 years ago (Liu et al. 2023b). This technique had spread to the Peiligang culture in the Central Plains by 8000 years ago (Liu et al. 2023a) and continuously practiced during the Yangshao culture (ca. 7000 − 5000 cal. BP) (Liu 2021). The brewing method observed at Pingliangtai clearly inherited this ancient tradition.
Husk phytoliths from foxtail millet, broomcorn millet, rice, and wheat were identified in the vessel residues. While it is conceivable that these husks could be associated with the use of sprouted grains for brewing beer, the presence of husk phytoliths in the fermented products may have resulted from deficient dehusking process. Hence, we are inclined to believe that the brewing method primarily involves qu fermentation.
4.2. Vessel functions and drinking behavior
According to previous research, jiandiping amphorae was the typical pottery types used as brewing and drinking vessels during the Yangshao culture period in the middle Yellow River region (Liu 2021). However, during the Longshan period, the vessels associated with alcohol consumption included various types of pottery serving multiple functions, ranging from production and consumption in multiple steps. The residue analysis in this study provides insights into the functions of specific vessel types. We categorize the pottery vessels into seven functional groups based on their form and associated microfossil remains (Fig. 6).
(1) Fermentation vessel. The guan jar (P13) features a restricted orifice and a globular body, a shape consistent with fermentation jars that first appeared in the early Neolithic (ca. 10,000-7000 cal. BP) across both southern and northern China (Liu et al. 2019). Over the following two millennia in North China, this vessel form disappeared in the Yanshao culture area of the western inland, but continued in the Beixin and Dawenkou cultures in the eastern coastal region (Liu 2023).
Starch granules inside the Pingliangtai jar exhibit gelatinization and fermentation-induced damage, along with phytoliths of broomcorn millet husks, as well as yeast cells and Monascus mold. The presence of these microfossils, combined with the vessel's form, suggests it was likely used for brewing and storing fermented beverages. This jar resembles corresponding vessels found in the Dawenkou and Longshan cultures in Shandong (Fig. 7:1,2).
2. Heating and pouring vessel. Gui pitchers (P9, P10, P11) have been regarded for heating and pouring out alcoholic beverages, as it is crafted with a spout, a handle, and three hollowed legs. In many cases, soot is visible on the exterior surface of the vessel. This vessel type was first developed in the east coast region and flourished in the Dawenkou and Shandong Longshan cultures (Tang 1979; Gao and Shao 1981). The Pingliangtai gui pitchers closely resemble those from the east coast region (Fig. 7: 3,4). Residue analysis on gui pitchers from the Liangchengzhen site of the Shandong Longshan culture indicates their association with fermented mixed beverages (McGovern et al. 2005). Starch granules exhibiting gelatinization and fermentation damage, along with fungal hyphae, yeast cells, and Monascus mold, were observed in the Pingliangtai gui pitchers. This confirms that the function of gui pitchers was related to preparing warm alcohol before serving.
3. Liquid serving vessels. The hu containers (P1, P3) are characterized by an open orifice and a cylindrical body with a globular curve. Diverse starch types showing characteristics of fermentation, yeast cells, and Monascus mold were present, suggesting their use for serving alcoholic beverages. The early examples of this vessel form can be traced to the Dawenkou and Shangdong Longshan cultures (Fig. 7:5,6).
4. Drinking, libation, and bartending vessels. The bei cup (P2) and gu goblet (P7) belong to this category. The bei cup closely resembles a modern drinking cup, with early forms found in the Dawenkou and Shandong Longshan cultures (Fig. 7:8,9). Residue analyses have revealed evidence of fermented beverages in such cups from Yuchisi (Dawenkou culture) and Liangchengzhen (McGovern et al. 2005; Liu et al. 2022).
The gu goblet features a tall cylindrical body and a flaring orifice, suggesting it was possibly used as a drinking cup. Its prototype can be found in the Dawenkou culture (Fig. 7:7). Some gu goblets have a very wide flaring orifice, raising questions about their practicality for drinking. However, our experimental drinking test with such a goblet indicates that these goblets can effectively serve as drinking vessels (Figure S3). The flaring orifice also suggests they may have been used for pouring liquids, not just for drinking. Similar bronze goblets with a bronze or jade rod placed inside, dating to the Shang and Zhou dynasties, have been found and were likely used as a set of bartending tools to facilitate liquid mixing for libation rituals (Li 2023). These libation rituals, which involve pouring alcohol onto the ground using gu goblets to serve deities, can be traced back to the Neolithic Liangzhu and Dawenkou cultures in the east coast region, predating the Longshan culture in Henan (Li 2022).
Starch granules inside the Pingliangtai cup and goblet show signs of fermentation damage, along with yeast cells and Monascus mold, suggesting they were used as drinking and libation vessels to serve the living and deities, respectively.
5. Cooking vessels. The ding cauldron (P5) and yan steamer (P12) were common cooking vessels. Ding was used for boiling food, while yan, with a grid base placed atop a cauldron, functioned as a steamer. Both vessels revealed gelatinized starches, consistent with their cooking functions. However, the yan steamer also contains starches showing characteristics of fermentation, yeast cells, and Monascus mold, suggesting that it was likely a multifunctional vessel used for steaming food, including ingredients used in fermentation. This finding is consistent with the residue analysis of a steamer rack from Liangchengzhen, which also revealed evidence of fermented beverages (McGovern et al. 2005). The Pingliangtai ding and yan also closely resemble those from Dawenkou and Longshan cultures in the east (Fig. 7:10–13).
6. Food serving vessel. The qigai lid (P4) contains diverse starches of almost all types, with some exhibiting signs of gelatinization and fermentation. Phytoliths from millet and wheat husks, as well as yeast cells, are also present. This lid was likely used to cover food vessels containing diverse foods, including fermented ones. Its form also suggests it may have served as a large bowl. This vessel form is widely distributed cross-regionally.
7. Processing vessel. The pen basin (P6) revealed a few starches from millets and snake gourd root, along with abundant phytoliths including millet husks and plant leaves, but no fungal particles were observed. This vessel was likely used for washing and processing plant foods, and also share similarities with those found in the Dawenkou culture (Fig. 7:14).
In summary, among the seven functional vessel types, four are specifically designated for alcohol production and consumption, including the guan jar, gui pitcher, hu container, bei cup, and gu goblet. Additionally, two types, the yan steamer and qigai lid, are multi-functional, also involving the processing of fermented foods. Together, the majority of common food vessel types at Pingliangtai are associated with alcohol-related consumption. This vessel assemblage represents a drinking behavior focusing on sequentially stepped activities, serving alcohol to individuals and deities. Through this process, individualized social status and hierarchy could be promoted and established. Such a sophisticated ritual system was likely related to feasting activities, which differ sharply from those of the previous Yangshao culture. Yangshao amphorae, along with funnel and stove-cauldron set, were used as the primary fermentation and drinking facilities (Fig. 7:15–17), and alcoholic beverages were shared by group drinking through straws, a behavior that emphasized simultaneous, communal activities with less focus on individual social status.
4.3. Regional interactions and emergence of new forms of ritual feasting in the Central Plains
The presence of diverse alcohol-related vessels reflects the development of feasting activities emphasizing personal social status in ceremonial contexts, a practice not rooted in the local Yangshao culture but likely originating in the Dawenkou culture in the east and later spreading to the Central Plains (Liu 2023). Archaeological data, particularly pottery types, indicate that the westward influence of the Dawenkou culture began around 3300 BC via the Ying River and its tributaries. The resulting material assemblages primarily belong to the Dawenkou culture but also include elements of the local Yangshao culture, together referred to as the Ying River variant of the Dawenkou culture (Du 1992).
This cultural diffusion was possibly facilitated by population migration, with specific routes starting along the Ying and Guo Rivers to eastern Henan, then continuously moving to central and western Henan. Another route possibly along the Huai River to southern Henan. Consequently, most Dawenkou cultural remains are found in eastern Henan, followed by central Henan, with fewer in other regions (Zhang 2003). Meanwhile, the Qujialing culture (3400 − 2500 BC) in the south also influenced the Central Plains (Meng 2011) (Fig. 7).
As a result, between 3300–2800 BC, the Central Plains experienced the coexistence and integration of the local tradition (late Yangshao culture) with non-indigenous traditions (the Dawenkou and Qujialing cultures). These changes are reflected in ceramic vessel forms. At Dahecun in Zhengzhou, for example, while traditional Yangshao style vessels (e.g., ding cauldrons, guan jars, bo bowls, pen basins, and jiandiping amphorae) were present, diverse new vessel forms from the Dawenkou culture (e.g., gui and he pitchers, hu jars, gaobingbei high-stand goblets) and Qujialing culture (e.g., dou high-stand plates, quanzubei ring-based cups) also entered the pottery assemblage.
By 2800–2400 BC, eastern Henan was predominantly under the influence of the Yuchisi variant of the late Dawenkou culture, which also displayed many elements of the early Longshan culture (Miaodigou II culture) originating in the Central Plains. The subsequent development of these mixtures of material culture from the east and west contributed to the formation of the Zaolütai variant of the Longshan culture, with Pingliangtai as the regional center (Luan 1997).
In the early period of the Zaolütai variant (2300 BC), influence from the eastern region once again became dominant in the Central Plains. Pottery types originating from the Shandong Longshan culture became predominant over the local Longshan variants.
During Pingliangtai's heyday (2200–2000 BC), many common vessel types, including those analyzed in this study (e.g., gui pitcher, hu goblet, gu container), appear to have originated in the Shandong Longshan culture. They have also been found at other regional centers or large sites along the Ying and Ru River regions, such as Wadian (Henan Provincial Institute of Cultural Relics and Archaeology 2004), Haojiatai (Henan Provincial Institute of Cultural Relics and Archaeology 2012), Yanzhai (Cultural relics and museums of Henan University and Xuchang Institute of Cultural Relics and Archaeology 2024), Wangchenggang (Henan Institute of Cultural Relics 1992), and Lutaigang (Zhengzhou University 2000). The prototypes of these vessels can be further traced back to Dawenkou culture sites, such as Yuchisi (Institute of Archaeology and Cultural Bureau of Mengcheng County 2007), Xixiahou (Gao and Ren 1964), Liulin (Nanjing Museum 1965), and Jianxin (He and Liu 1996) (Fig. 7).
Analysis of pottery residues from Pingliangtai in this study further demonstrates that the majority of vessel types constitute a complex assemblage used for serving alcoholic drinks in ritual feasts, and this assemblage was dispersed from the east to the Central Plains. This cultural phenomenon aligns with ancient texts documenting interactions and integration between the Huaxia groups in the west and the Dongyi groups in the east (Li 1983; Jin 2006).
These alcohol-related vessels facilitated individual-centered feasting behavior, introducing new cultural values and encouraging social hierarchy. This shift parallels other social changes in the Longshan culture, such as the construction of walled enclosures for inter-group competition and the production and exchange of prestige goods as status symbols. Together, these developments set the foundation for the emergence of the Bronze Age state of the Erlitou culture.