Abushady H, Bashandy A, Aziz N, Ibrahim H (2005): Molecular characterization of Bacillus subtilis surfactin producing strain and the factors affecting its production. International Journal of Agriculture and Biology 3, 337-344
Al-Bahry S, Al-Wahaibi Y, Elshafie A, Al-Bemani A, Joshi S, Al-Makhmari H, Al-Sulaimani H (2013): Biosurfactant production by Bacillus subtilis B20 using date molasses and its possible application in enhanced oil recovery. International Biodeterioration & Biodegradation 81, 141-146
Al-Dhabi NA, Esmail GA, Valan Arasu M (2020): Enhanced Production of Biosurfactant from Bacillus subtilis Strain Al-Dhabi-130 under Solid-State Fermentation Using Date Molasses from Saudi Arabia for Bioremediation of Crude-Oil-Contaminated Soils. International Journal of Environmental Research and Public Health 17, 8446
Anvari S, Hajfarajollah H, Mokhtarani B, Noghabi KA (2015): Physiochemical and thermodynamic characterization of lipopeptide biosurfactant secreted by Bacillus tequilensis HK01. RSC advances 5, 91836-91845
Balan SS, Kumar CG, Jayalakshmi S (2017): Aneurinifactin, a new lipopeptide biosurfactant produced by a marine Aneurinibacillus aneurinilyticus SBP-11 isolated from Gulf of Mannar: Purification, characterization and its biological evaluation. Microbiological research 194, 1-9
Bartal A, Vigneshwari A, Bóka B, Vörös M, Takács I, Kredics L, Manczinger L, Varga M, Vágvölgyi C, Szekeres A (2018): Effects of different cultivation parameters on the production of surfactin variants by a Bacillus subtilis strain. Molecules 23, 2675
Beltran-Gracia E, Macedo-Raygoza G, Villafaña-Rojas J, Martinez-Rodriguez A, Chavez-Castrillon YY, Espinosa-Escalante FM, Di Mascio P, Ogura T, Beltran-Garcia MJ (2017): Production of lipopeptides by fermentation processes: Endophytic bacteria, fermentation strategies and easy methods for bacterial selection. Fermentation processes, 199-222
Bertrand B, Martínez-Morales F, Rosas-Galván NS, Morales-Guzmán D, Trejo-Hernández MR (2018): Statistical design, a powerful tool for optimizing biosurfactant production: a review. Colloids and Interfaces 2, 36
Biniarz P, Łukaszewicz M, Janek T (2017): Screening concepts, characterization and structural analysis of microbial-derived bioactive lipopeptides: a review. Critical reviews in biotechnology 37, 393-410
Chandrasekaran M, Bahkali AH (2013): Valorization of date palm (Phoenix dactylifera) fruit processing by-products and wastes using bioprocess technology–Review. Saudi journal of biological sciences 20, 105-120
Chen Y, Liu S, Mou H, Ma Y, Li M, Hu X (2017): Characterization of lipopeptide biosurfactants produced by Bacillus licheniformis MB01 from marine sediments. Front. Microbiol. 2017, V. 8
Cho KM, Math RK, Hong SY, Islam SMA, Mandanna DK, Cho JJ, Yun MG, Kim JM, Yun HD (2009): Iturin produced by Bacillus pumilus HY1 from Korean soybean sauce (kanjang) inhibits growth of aflatoxin producing fungi. Food Control 20, 402-406
Chooklin CS, Phertmean S, Cheirsilp B, Maneerat S, Saimmai A (2013): Utilization of palm oil mill effluent as a novel and promising substrate for biosurfactant production by Nevskia ramosa NA3. Songklanakarin Journal of Science & Technology 35
Coronel-León J, de Grau G, Grau-Campistany A, Farfan M, Rabanal F, Manresa A, Marqués AM (2015): Biosurfactant production by AL 1.1, a Bacillus licheniformis strain isolated from Antarctica: production, chemical characterization and properties. Annals of microbiology 65, 2065-2078
Dang Y, Zhao F, Liu X, Fan X, Huang R, Gao W, Wang S, Yang C (2019): Enhanced production of antifungal lipopeptide iturin A by Bacillus amyloliquefaciens LL3 through metabolic engineering and culture conditions optimization. Microbial cell factories 18, 1-14
Eswari JS, Anand M, Venkateswarlu C (2016): Optimum culture medium composition for lipopeptide production by Bacillus subtilis using response surface model-based ant colony optimization. Sadhana 41, 55-65
Feng J-Q, Gang H-Z, Li D-S, Liu J-F, Yang S-Z, Mu B-Z (2019): Characterization of biosurfactant lipopeptide and its performance evaluation for oil-spill remediation. RSC advances 9, 9629-9632
Ghasemi A, Moosavi-Nasab M, Behzadnia A, Rezaei M (2018): Enhanced biosurfactant production with low-quality date syrup by Lactobacillus rhamnosus using a fed-batch fermentation. Food science and biotechnology 27, 1137-1144
Ghazala I, Bouassida M, Krichen F, Manuel Benito J, Ellouz‐Chaabouni S, Haddar A (2017): Anionic lipopeptides from Bacillus mojavensis I4 as effective antihypertensive agents: Production, characterization, and identification. Engineering in life sciences 17, 1244-1253
Ghazala I, Bouallegue A, Haddar A, Ellouz-Chaabouni S (2019): Characterization and production optimization of biosurfactants by Bacillus mojavensis I4 with biotechnological potential for microbial enhanced oil recovery. Biodegradation 30, 235-245
Ghribi D, Abdelkefi-Mesrati L, Mnif I, Kammoun R, Ayadi I, Saadaoui I, Maktouf S, Chaabouni-Ellouze S (2012): Investigation of antimicrobial activity and statistical optimization of Bacillus subtilis SPB1 biosurfactant production in solid-state fermentation. Journal of Biomedicine and Biotechnology 2012
Habib S, Ahmad SA, Wan Johari WL, Abd Shukor MY, Alias SA, Smykla J, Saruni NH, Abdul Razak NS, Yasid NA (2020): Production of lipopeptide biosurfactant by a hydrocarbon-degrading Antarctic Rhodococcus. International journal of molecular sciences 21, 6138
Hippolyte MT, Augustin M, Hervé TM, Robert N, Devappa S (2018): Application of response surface methodology to improve the production of antimicrobial biosurfactants by Lactobacillus paracasei subsp. tolerans N2 using sugar cane molasses as substrate. Bioresources and Bioprocessing 5, 1-16
Hmidet N, Ben Ayed H, Jacques P, Nasri M (2017): Enhancement of surfactin and fengycin production by Bacillus mojavensis A21: application for diesel biodegradation. BioMed research international 2017
Hu F, Liu Y, Li S (2019): Rational strain improvement for surfactin production: enhancing the yield and generating novel structures. Microbial cell factories 18, 1-13
Huszcza E, Burczyk B (2003): Biosurfactant production by Bacillus coagulans. Journal of Surfactants and Detergents 6, 61-64
Ismail W, Al-Rowaihi IS, Al-Humam AA, Hamza RY, El Nayal AM, Bououdina M (2013): Characterization of a lipopeptide biosurfactant produced by a crude-oil-emulsifying Bacillus sp. I-15. International Biodeterioration & Biodegradation 84, 168-178
Jemil N, Ayed HB, Manresa A, Nasri M, Hmidet N (2017): Antioxidant properties, antimicrobial and anti-adhesive activities of DCS1 lipopeptides from Bacillus methylotrophicus DCS1. BMC microbiology 17, 1-11
Joshi S, Bharucha C, Jha S, Yadav S, Nerurkar A, Desai AJ (2008): Biosurfactant production using molasses and whey under thermophilic conditions. Bioresource technology 99, 195-199
Joy S, Rahman PK, Sharma S (2017): Biosurfactant production and concomitant hydrocarbon degradation potentials of bacteria isolated from extreme and hydrocarbon contaminated environments. Chemical Engineering Journal 317, 232-241
Kanna R, Gummadi SN, Kumar GS (2014): Production and characterization of biosurfactant by Pseudomonas putida MTCC 2467. Journal of Biological Sciences 14, 436
Kebbouche-Gana S, Gana ML, Ferrioune I, Khemili S, Lenchi N, Akmouci-Toumi S, Bouanane-Darenfed NA, Djelali N-E (2013): Production of biosurfactant on crude date syrup under saline conditions by entrapped cells of Natrialba sp. strain E21, an extremely halophilic bacterium isolated from a solar saltern (Ain Salah, Algeria). Extremophiles 17, 981-993
Kim KM, Lee JY, Kim CK, Kang JS (2009): Isolation and characterization of surfactin produced by Bacillus polyfermenticus KJS-2. Archives of pharmacal research 32, 711-715
Kiran GS, Priyadharsini S, Sajayan A, Priyadharsini GB, Poulose N, Selvin J (2017): Production of lipopeptide biosurfactant by a marine Nesterenkonia sp. and its application in food industry. Frontiers in microbiology 8, 1138
Liang X, Shi R, Radosevich M, Zhao F, Zhang Y, Han S, Zhang Y (2017): Anaerobic lipopeptide biosurfactant production by an engineered bacterial strain for in situ microbial enhanced oil recovery. RSC advances 7, 20667-20676
Lin H-Y, Rao YK, Wu W-S, Tzeng Y-M (2007): Ferrous ion enhanced lipopeptide antibiotic iturin A production from Bacillus amyloliquefaciens B128. International Journal of Applied Science and Engineering 5, 123-132
Liu X, Ren B, Gao H, Liu M, Dai H, Song F, Yu Z, Wang S, Hu J, Kokare CR (2012): Optimization for the production of surfactin with a new synergistic antifungal activity. PloS one 7, e34430
Luna JM, Rufino RD, Jara AMA, Brasileiro PP, Sarubbo LA (2015): Environmental applications of the biosurfactant produced by Candida sphaerica cultivated in low-cost substrates. Colloids and Surfaces A: Physicochemical and Engineering Aspects 480, 413-418
Mani P, Sivakumar P, Balan SS (2016): Economic production and oil recovery efficiency of a lipopeptide biosurfactant from a novel marine bacterium Bacillus simplex. Achievements in the Life Sciences 10, 102-110
Martins PC, Bastos CG, Granjeiro PA, Martins VG (2018): New lipopeptide produced by Corynebacterium aquaticum from a low-cost substrate. Bioprocess and biosystems engineering 41, 1177-1183
Mnif I, Sahnoun R, Ellouze-Chaabouni S, Ghribi D (2013a): Evaluation of B. subtilis SPB1 biosurfactants' potency for diesel-contaminated soil washing: optimization of oil desorption using Taguchi design. Environmental Science and Pollution Research 21, 851-861
Mnif I, Ellouze‐Chaabouni S, Ghribi D (2013b): Economic production of Bacillus subtilis SPB1 biosurfactant using local agro‐industrial wastes and its application in enhancing solubility of diesel. Journal of Chemical Technology & Biotechnology 88, 779-787
Mnif I, Elleuch M, Chaabouni SE, Ghribi D (2013c): Bacillus subtilis SPB1 biosurfactant: production optimization and insecticidal activity against the carob moth Ectomyelois ceratoniae. Crop Protection 50, 66-72
Mnif I, Besbes S, Ellouze‐Ghorbel R, Ellouze‐Chaabouni S, Ghribi D (2013d): Improvement of bread dough quality by Bacillus subtilis SPB1 biosurfactant addition: optimized extraction using response surface methodology. Journal of the Science of Food and Agriculture 93, 3055-3064
Mnif I, Ellouze‐Chaabouni S, Ayedi Y, Ghribi D (2014): Treatment of diesel‐and kerosene‐contaminated water by B. subtilis SPB1 biosurfactant‐producing strain. Water Environment Research 86, 707-716
Mnif I, Ghribi D (2015a-a): Microbial derived surface active compounds: properties and screening concept. World Journal of Microbiology and Biotechnology 31, 1001-1020
Mnif I, Ghribi D (2015a-b): Microbial derived surface active compounds: properties and screening concept. World Journal of Microbiology Biotechnology 31, 1001-1020
Mnif I, Mnif S, Sahnoun R, Maktouf S, Ayedi Y, Ellouze-Chaabouni S, Ghribi D (2015a): Biodegradation of diesel oil by a novel microbial consortium: comparison between co-inoculation with biosurfactant-producing strain and exogenously added biosurfactants. Environmental Science and Pollution Research 22, 14852-14861
Mnif I, Fendri R, Ghribi D (2015b): Malachite green bioremoval by a newly isolated strain Citrobacter sedlakii RI11; enhancement of the treatment by biosurfactant addition. Water Science and Technology 72, 1283-1293
Mnif I, Ghribi D (2015b): Lipopeptide surfactants: production, recovery and pore forming capacity. Peptides 71, 100-112
Mnif I, Fendri R, Ghribi D (2015c): Biosorption of Congo Red from aqueous solution by Bacillus weihenstephanensis RI12; effect of SPB1 biosurfactant addition on biodecolorization potency. Water Science and Technology 72, 865-874
Mnif I, Ghribi D (2015c): Review lipopeptides biosurfactants: mean classes and new insights for industrial, biomedical, and environmental applications. Peptide Science 104, 129-147
Mnif I, Maktouf S, Fendri R, Kriaa M, Ellouze S, Ghribi D (2016a): Improvement of methyl orange dye biotreatment by a novel isolated strain, Aeromonas veronii GRI, by SPB1 biosurfactant addition. Environmental Science and Pollution Research 23, 1742-1754
Mnif I, Grau-Campistany A, Coronel-León J, Hammami I, Triki MA, Manresa A, Ghribi D (2016b): Purification and identification of Bacillus subtilis SPB1 lipopeptide biosurfactant exhibiting antifungal activity against Rhizoctonia bataticola and Rhizoctonia solani. Environmental Science and Pollution Research 23, 6690-6699
Moshtagh B, Hawboldt K, Zhang B (2019): Optimization of biosurfactant production by Bacillus subtilis N3-1P using the brewery waste as the carbon source. Environmental technology 40, 3371-3380
Mukherjee S, Das P, Sivapathasekaran C, Sen R (2009): Antimicrobial biosurfactants from marine Bacillus circulans: extracellular synthesis and purification. Letters in applied microbiology 48, 281-288
Najib H, Al‐Yousef YM (2013): Value‐Added Utilization of Dates By‐Products. Dates: Postharvest Science, Processing Technology and Health Benefits, 233-260
Nancib N, Nancib A, Boudjelal A, Benslimane C, Blanchard F, Boudrant J (2001): The effect of supplementation by different nitrogen sources on the production of lactic acid from date juice by Lactobacillus casei subsp. rhamnosus. Bioresource Technology 78, 149-153
Ndlovu T, Rautenbach M, Khan S, Khan W (2017): Variants of lipopeptides and glycolipids produced by Bacillus amyloliquefaciens and Pseudomonas aeruginosa cultured in different carbon substrates. AMB Express 7, 1-13
Nurfarahin AH, Mohamed MS, Phang LY (2018): Culture medium development for microbial-derived surfactants production—an overview. Molecules 23, 1049
Ohadi M, Dehghannoudeh G, Shakibaie M, Banat IM, Pournamdari M, Forootanfar H (2017): Isolation, characterization, and optimization of biosurfactant production by an oil-degrading Acinetobacter junii B6 isolated from an Iranian oil excavation site. Biocatalysis and Agricultural Biotechnology 12, 1-9
Olasanmi IO, Thring RW (2018): The role of biosurfactants in the continued drive for environmental sustainability. Sustainability 10, 4817
Pacwa-Płociniczak M, Płaza GA, Piotrowska-Seget Z, Cameotra SS (2011): Environmental applications of biosurfactants: recent advances. International Journal of Molecular Sciences 12, 633-654
Paraszkiewicz K, Bernat P, Kuśmierska A, Chojniak J, Płaza G (2018): Structural identification of lipopeptide biosurfactants produced by Bacillus subtilis strains grown on the media obtained from renewable natural resources. Journal of Environmental Management 209, 65-70
Pathak KV, Keharia H (2014): Application of extracellular lipopeptide biosurfactant produced by endophytic Bacillus subtilis K1 isolated from aerial roots of banyan (Ficus benghalensis) in microbially enhanced oil recovery (MEOR). 3 Biotech 4, 41-48
Patowary K, Patowary R, Kalita MC, Deka S (2017): Characterization of biosurfactant produced during degradation of hydrocarbons using crude oil as sole source of carbon. Frontiers in microbiology 8, 279
Pereira JF, Gudiña EJ, Costa R, Vitorino R, Teixeira JA, Coutinho JA, Rodrigues LR (2013): Optimization and characterization of biosurfactant production by Bacillus subtilis isolates towards microbial enhanced oil recovery applications. Fuel 111, 259-268
Phulpoto IA, Yu Z, Hu B, Wang Y, Ndayisenga F, Li J, Liang H, Qazi MA (2020): Production and characterization of surfactin-like biosurfactant produced by novel strain Bacillus nealsonii S2MT and it's potential for oil contaminated soil remediation. Microbial cell factories 19, 1-12
Pueyo MT, Bloch C, Carmona-Ribeiro AM, Di Mascio P (2009): Lipopeptides produced by a soil Bacillus megaterium strain. Microbial ecology 57, 367
Purwasena IA, Astuti DI, Syukron M, Amaniyah M, Sugai Y (2019): Stability test of biosurfactant produced by Bacillus licheniformis DS1 using experimental design and its application for MEOR. Journal of Petroleum Science and Engineering 183, 106383
Qiao N, Shao Z (2010): Isolation and characterization of a novel biosurfactant produced by hydrocarbon‐degrading bacterium Alcanivorax dieselolei B‐5. Journal of applied microbiology 108, 1207-1216
Ramírez IM, Tsaousi K, Rudden M, Marchant R, Alameda EJ, Román MG, Banat IM (2015): Rhamnolipid and surfactin production from olive oil mill waste as sole carbon source. Bioresource technology 198, 231-236
Rodrigues L, Teixeira J, Oliveira R (2006): Low-cost fermentative medium for biosurfactant production by probiotic bacteria. Biochemical Engineering Journal 32, 135-142
Roy A, Mahata D, Paul D, Korpole S, Franco OL, Mandal SM (2013): Purification, biochemical characterization and self-assembled structure of a fengycin-like antifungal peptide from Bacillus thuringiensis strain SM1. Frontiers in microbiology 4, 332
Sarubbo LA, Luna J, Rufino RD, Brasileiro P (2016): Production of a Low-cost Biosurfactant for Application in the Remediation of Sea water Contaminated with Petroleum Derivates. Chemical Engineering Transactions 49, 523-528
Shi J, Zhu X, Lu Y, Zhao H, Lu F, Lu Z (2018): Improving iturin A production of Bacillus amyloliquefaciens by genome shuffling and its inhibition against Saccharomyces cerevisiae in orange juice. Frontiers in microbiology 9, 2683
Shih L, Kuo C-Y, Hsieh F-C, Kao S-S, Hsieh C (2008): Use of surface response methodology to optimize culture conditions for iturin A production by Bacillus subtilis in solid-state fermentation. Journal of the Chinese Institute of Chemical Engineers 39, 635-643
Singh AK, Rautela R, Cameotra SS (2014): Substrate dependent in vitro antifungal activity of Bacillus sp strain AR2. Microbial cell factories 13, 1-11
Singh P, Patil Y, Rale V (2019): Biosurfactant production: emerging trends and promising strategies. Journal of applied microbiology 126, 2-13
Soares CC, de Andrade ASA, Ferreira GFD, de Almeida AF, Druzian JI, Lobato AKdCL (2018): Biosurfactant production by Bacillus subtilis UFPEDA 86 using papaya (Carica papaya L.) waste as substrate: Viability studies and pH influence of the culture medium. African Journal of Biotechnology 17, 928-937
Takashi A, Jin GY, Mizumoto S, Shahedur RM, Okuno K, Shoda M (2009): Solid state fermentation of lipopeptide antibiotic iturin A by using a novel solid state fermentation reactor system. Journal of Environmental Sciences 21, S162-S165
Thimon L, Peypoux F, Michel G (1992): Interactions of surfactin, a biosurfactant from Bacillus subtilis, with inorganic cations. Biotechnology letters 14, 713-718
Varadavenkatesan T, Murty VR (2013): Production of a lipopeptide biosurfactant by a novel Bacillus sp. and its applicability to enhanced oil recovery. International Scholarly Research Notices 2013
Wang Q, Chen S, Zhang J, Sun M, Liu Z, Yu Z (2008): Co-producing lipopeptides and poly-γ-glutamic acid by solid-state fermentation of Bacillus subtilis using soybean and sweet potato residues and its biocontrol and fertilizer synergistic effects. Bioresource technology 99, 3318-3323
Wei Y-H, Chu I-M (2002): Mn 2+ improves surfactin production by Bacillus subtilis. Biotechnology Letters 24, 479-482
Wei YH, Wang LF, Chang JS (2004): Optimizing iron supplement strategies for enhanced surfactin production with Bacillus subtilis. Biotechnology Progress 20, 979-983
Yadav AK, Bipinraj NK, Chaudhari AB, Kothari RM (2011): Production of L (+) lactic acid from sweet sorghum, date palm, and golden syrup as alternative carbon sources. Starch‐Stärke 63, 632-636
Zhang J, Xue Q, Gao H, Lai H, Wang P (2016): Production of lipopeptide biosurfactants by Bacillus atrophaeus 5-2a and their potential use in microbial enhanced oil recovery. Microbial cell factories 15, 1-11
Zouari O, Lecouturier D, Rochex A, Chataigne G, Dhulster P, Jacques P, Ghribi D (2019b): Bio-emulsifying and biodegradation activities of syringafactin producing Pseudomonas spp. strains isolated from oil contaminated soils. Biodegradation 30, 259-272
Zouari R, Ellouze-Chaabouni S, Ghribi-Aydi D (2014): Optimization of Bacillus subtilis SPB1 biosurfactant production under solid-state fermentation using by-products of a traditional olive mill factory. Achievements in the Life Sciences 8, 162-169
Zouari R, Ellouze-Chaabouni S, Ghribi D (2019a): Use of Butter Milk and Poultry-Transforming Wastes for Enhanced Production of Bacillus subtilis SPB1 Biosurfactant in Submerged Fermentation. Journal of Microbiology, Biotechnology and Food Sciences 2019b, 462-466