1.Rushton PJ, Somssich IE, Ringler P, Shen QXJ: WRKY transcription factors. Trends Plant Sci. 2010, 15(5):247–258.
2.Zhang Y, Wang L: The WRKY transcription factor superfamily: its origin in eukaryotes and expansion in plants. BMC Evol Biol. 2005, 5(1):1.
3.Chen L, Xiang S, Chen Y, Li D, Yu D: Arabidopsis WRKY45 Interacts with the DELLA Protein RGL1 to Positively Regulate Age-Triggered Leaf Senescence. Mol Plant 2017, 10(9):1174–1189.
4.Tian X, Li X, Zhou W, Ren Y, Wang Z, Liu Z, Tang J, Tong H, Fang J, Bu Q: Transcription Factor OsWRKY53 Positively Regulates Brassinosteroid Signaling and Plant Architecture. Plant Phys. 2017, 175(3):1337–1349.
5.Hsu FC, Chou MY, Chou SJ, Li YR, Peng HP, Shih MC: Submergence confers immunity mediated by the WRKY22 transcription factor in Arabidopsis. Plant Cell 2013, 25(7):2699–2713.
6.Hu Y, Chen L, Wang H, Zhang L, Wang F, Yu D: Arabidopsis transcription factor WRKY8 functions antagonistically with its interacting partner VQ9 to modulate salinity stress tolerance. Plant J. 2013, 74(5):730–745.
7.Mao G, Meng X, Liu Y, Zheng Z, Chen Z, Zhang S: Phosphorylation of a WRKY transcription factor by two pathogen-responsive MAPKs drives phytoalexin biosynthesis in Arabidopsis. Plant Cell 2011, 23(4):1639–1653.
8.Kim KC, Lai ZB, Fan BF, Chen ZX: Arabidopsis WRKY38 and WRKY62 Transcription Factors Interact with Histone Deacetylase 19 in Basal Defense. Plant Cell 2008, 20(9):2357–2371.
9.Qi W, Chen X, Fang P, Shi S, Li J, Liu X, Cao X, Zhao N, Hao H, Li Y et al: Genomic and transcriptomic sequencing of Rosa hybrida provides microsatellite markers for breeding, flower trait improvement and taxonomy studies. BMC plant biology 2018, 18(1):119.
10.Liu X, Cao X, Shi S, Zhao N, Li D, Fang P, Chen X, Qi W, Zhang Z: Comparative RNA-Seq analysis reveals a critical role for brassinosteroids in rose (Rosa hybrida) petal defense against Botrytis cinerea infection. BMC genetics 2018, 19(1):62.
11.Hao Y, Cao X, Ma C, Zhang Z, Zhao N, Ali A, Hou T, Xiang Z, Zhuang J, Wu S et al: Potential Applications and Antifungal Activities of Engineered Nanomaterials against Gray Mold Disease Agent Botrytis cinerea on Rose Petals. Frontiers in Plant Science 2017, 8(1332).
12.Wang Y, Tang H, Debarry JD, Tan X, Li J, Wang X, Lee TH, Jin H, Marler B, Guo H et al: MCScanX: a toolkit for detection and evolutionary analysis of gene synteny and collinearity. Nucleic acids research 2012, 40(7):e49.
13.Gao FL, Chen CJ, Arab DA, Du ZG, He YH, Ho SYW: EasyCodeML: A visual tool for analysis of selection using CodeML. Ecol Evol 2019, 9(7):3891–3898.
14.Liu YL, Schiff M, Dinesh-Kumar SP: Virus-induced gene silencing in tomato. The Plant journal 2002, 31(6):777–786.
15.Raymond O, Gouzy J, Just J, Badouin H, Verdenaud M, Lemainque A, Vergne P, Moja S, Choisne N, Pont C et al: The Rosa genome provides new insights into the domestication of modern roses. Nature genetics 2018.
16.Wang Q, Wang M, Zhang X, Hao B, Kaushik SK, Pan Y: WRKY gene family evolution in Arabidopsis thaliana. Genetica 2011, 139(8):973–983.
17.Ross CA, Liu Y, Shen QXJ: The WRKY gene family in rice (Oryza sativa). J Integr Plant Biol 2007, 49(6):827–842.
18.Huang SX, Gao YF, Liu JK, Peng XL, Niu XL, Fei ZJ, Cao SQ, Liu YS: Genome-wide analysis of WRKY transcription factors in Solanum lycopersicum. Mol Genet Genomics 2012, 287(6):495–513.
19.Dou L, Zhang X, Pang C, Song M, Wei H, Fan S, Yu S: Genome-wide analysis of the WRKY gene family in cotton. Mol Genet Genomics 2014, 289(6):1103–1121.
20.Ling J, Jiang W, Zhang Y, Yu H, Mao Z, Gu X, Huang S, Xie B: Genome-wide analysis of WRKY gene family in Cucumis sativus. BMC genomics 2011, 12:471.
21.He H, Dong Q, Shao Y, Jiang H, Zhu S, Cheng B, Xiang Y: Genome-wide survey and characterization of the WRKY gene family in Populus trichocarpa. Plant Cell Rep 2012, 31(7):1199–1217.
22.Mondragon-Palomino M, Meyers BC, Michelmore RW, Gaut BS: Patterns of positive selection in the complete NBS-LRR gene family of Arabidopsis thaliana. Genome Res 2002, 12(9):1305–1315.
23.Lai Z, Vinod KM, Zheng Z, Fan B, Chen Z: Roles of ArabidopsisWRKY3 and WRKY4 Transcription Factors in Plant Responses to Pathogens. BMC plant biology 2008, 8(1):68.
24.Kim K-C, Baofang F, Zhixiang C: Pathogen-induced Arabidopsis WRKY7 is a transcriptional repressor and enhances plant susceptibility to Pseudomonas syringae. Plant Physiology 2006, 142(3):1180–1192.
25.Chen L, Zhang L, Yu D: Wounding-Induced WRKY8 Is Involved in Basal Defense in\\r <i>Arabidopsis</i>. Mol Plant Microbe Interact 2010, 23(5):558–565.
26.Jiang CH, Huang ZY, Xie P, Gu C, Li K, Wang DC, Yu YY, Fan ZH, Wang CJ, Wang YP: Transcription factors WRKY70 and WRKY11 served as regulators in rhizobacterium Bacillus cereus AR156-induced systemic resistance to Pseudomonas syringae pv. tomato DC3000 in Arabidopsis. Proceedings of the Prehistoric Society 2016, 41(1):156–253.
27.Journot Catalino NL, Somssich IE, Dominique R, Thomas K: The transcription factors WRKY11 and WRKY17 act as negative regulators of basal resistance in Arabidopsis thaliana. Plant Cell 2006, 18(11):3289–3302.
28.Xu X, Chen C, Fan B, Chen Z: Physical and functional interactions between pathogen-induced Arabidopsis WRKY18, WRKY40, and WRKY60 transcription factors. Plant Cell 2006, 18(5):1310–1326.
29.Zheng Z, Mosher SL, Fan B, Klessig DF, Chen Z: Functional analysis of Arabidopsis WRKY25 transcription factor in plant defense against Pseudomonas syringae. BMC Plant Biology,7,1(2007–01–10) 2007, 7(1):2–2.
30.Mukhtar MS, Marco Y, Somssich IE: The Arabidopsis transcription factor WRKY27 influences wilt disease symptom development caused by Ralstonia solanacearum. Plant Journal for Cell & Molecular Biology 2010, 56(6):935–947.
31.Chen X, Chen, Jun L, Guifang L, Airong W, Zonghua W, Guodong L: Overexpression of AtWRKY28 and AtWRKY75 in Arabidopsis enhances resistance to oxalic acid and Sclerotinia sclerotiorum. Plant Cell Reports 2013, 32(10):1589–1599.
32.Zheng Z, Qamar SA, Chen Z, Mengiste T: Arabidopsis WRKY33 transcription factor is required for resistance to necrotrophic fungal pathogens. The Plant Journal 2006, 48(4):592–605.
33.Kim K-C, Zhibing L, Baofang F, Zhixiang C: Arabidopsis WRKY38 and WRKY62 transcription factors interact with histone deacetylase 19 in basal defense. Plant Cell 2008, 20(9):2357–2371.
34.Hu Y, Dong Q, Yu D: Arabidopsis WRKY46 coordinates with WRKY70 and WRKY53 in basal resistance against pathogen Pseudomonas syringae. Plant Science 2012, 185–186(x):288–297.
35.Xing D-H, Zi-Bing, Zu-Yu, Zheng, Vinod, Bao-Fang, Zhi-Xiang, Chen: Stress- and Pathogen-Induced Arabidopsis WRKY48 is a Transcriptional Activator that Represses Plant Basal Defense. Molecular Plant 2008, 1(3):459–470.
36.Gao Q-M, Srivathsa V, Duroy N, Aardra K: Low oleic acid-derived repression of jasmonic acid-inducible defense responses requires the WRKY50 and WRKY51 proteins. Plant Physiology 2011, 155(1):464–476.
37.Murray SL, Ingle RA, Petersen LN, Denby KJ: Basal Resistance Against\\r <i>Pseudomonas syringae</i>\\r in\\r <i>Arabidopsis</i>\\r Involves WRKY53 and a Protein with Homology to a Nematode Resistance Protein. Mol Plant Microbe Interact 2007, 20(11):1431–1438.
38.Jiang Y, Yu D: The WRKY57 Transcription Factor Affects the Expression of Jasmonate ZIM-Domain Genes Transcriptionally to Compromise Botrytis cinerea Resistance. Plant Physiology 2016, 171(4):2771–2782.
39.Gao R, Liu P, Yong Y, Wong SM: Genome-wide transcriptomic analysis reveals correlation between higher WRKY61 expression and reduced symptom severity in Turnip crinkle virus infectedArabidopsis thaliana. Scientific Reports 2016, 6(1):24604.
40.Liu X, Song Y, Xing F, Wang N, Wen F, Zhu C: GhWRKY25, a group I WRKY gene from cotton, confers differential tolerance to abiotic and biotic stresses in transgenic Nicotiana benthamiana. Protoplasma 2015, 253(5):1265–1281.
41.Guo R, Qiao H, Zhao J, Wang X, Wang X: The Grape VlWRKY3 Gene Promotes Abiotic and Biotic Stress Tolerance in Transgenic Arabidopsis thaliana. Frontiers in Plant Science 2018, 9:545-.
42.Merz PR, Moser T, Höll J, Kortekamp A, Buchholz G, Zyprian E, Bogs J: The transcription factor VvWRKY33 is involved in the regulation of grapevine (Vitis vinifera) defense against the oomycete pathogen Plasmopara viticola. Physiologia Plantarum 2014, 153(3):365.
43.Adachi H, Ishihama N, Nakano T, Yoshioka M, Yoshioka H: Nicotiana benthamiana MAPK-WRKY pathway confers resistance to a necrotrophic pathogen Botrytis cinerea. Plant Signal Behav 2016, 11(6):e1183085.
44.Han M, Ryu HS, Kim CY, Park DS, Ahn YK, Jeon JS: OsWRKY30 is a transcription activator that enhances rice resistance to the Xanthomonas oryzae pathovar oryzae. Journal of Plant Biology 2013, 56(4):258–265.
45.Peng X, Hu Y, Tang X, Zhou P, Deng X, Wang H, Guo Z: Constitutive expression of rice WRKY30 gene increases the endogenous jasmonic acid accumulation, PR gene expression and resistance to fungal pathogens in rice. Planta 2012, 236(5):1485–1498.
46.Chujo T, Koji M, Satoshi O, Yuka M, Takafumi S, Mitsuko KK, Akira T, Yoko N, Eiichi M, Hideaki N: Overexpression of phosphomimic mutated OsWRKY53 leads to enhanced blast resistance in rice. Plos One 2014, 9(6):e98737.
47.Liu X, Xian, Quan, Qian, QIAN, WANG, Ming, Sheng, CHEN: OsWRKY03, a rice transcriptional activator that functions in defense signaling pathway upstream of OsNPR1. Cell Research 2005, 15(8):593–603.
48.Wang H, Meng J, Peng X, Tang X, Zhou P, Xiang J, Deng X: Rice WRKY4 acts as a transcriptional activator mediating defense responses toward Rhizoctonia solani, the causing agent of rice sheath blight. Plant molecular biology 2015, 89(1–2):157–171.
49.Abbruscato P, Nepusz T, Mizzi L, Del CM, Morandini P, Fumasoni I, Michel C, Paccanaro A, Guiderdoni E, Schaffrath U: OsWRKY22, a monocot WRKY gene, plays a role in the resistance response to blast. Molecular Plant Pathology 2012, 13(8):828–841.
50.Liu X, Bai X, Wang X, Chu C: OsWRKY71, a rice transcription factor, is involved in rice defense response. Journal of Plant Physiology 2007, 164(8):969–979.
51.Zhang J, Youliang, Peng, Zejian: Constitutive expression of pathogen-inducible OsWRKY31 enhances disease resistance and affects root growth and auxin response in transgenic rice plants. Cell Research 2008, 18(4):508–521.
52.Ellis JG, Dodds PN, Lawrence GJ: The role of secreted proteins in diseases of plants caused by rust, powdery mildew and smut fungi. Current Opinion in Microbiology 2007, 10(4):326–331.
53.Yang B, Jiang Y, Rahman MH, Deyholos MK, Kav NN: Identification and expression analysis of WRKY transcription factor genes in canola (Brassica napus L.) in response to fungal pathogens and hormone treatments. BMC plant biology 2009, 9(1):68.
54.Oh S-K, Kwang-Hyun B, Jeong Mee P, So Young Y, Seung Hun Y, Sophien K, Doil C: Capsicum annuum WRKY protein CaWRKY1 is a negative regulator of pathogen defense. New Phytologist 2010, 177(4):977–989.